Professor and Chair

University of Rochester •

01/1982 – 01/2000 •

Pharmacology and Physiology

Associate Professor & Professor

University of Minnesota System •

01/1969 – 01/1982 •

Pharmacology

Assistant Professor/Associate Professort

Cornell University •

01/1965 – 01/1969 •

Physiology, Pharmacology, and Biochemistry

University of Wisconsin–Madison •

01/1960 – 01/1962 •

Pharmacology

D.V.M.

Iowa State University •

01/1954 – 01/1960 •

Veterinary Medicine